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Which wires are you confused about?  The red and black tangle of wires in the 3rd picture appear to be the center roller and frame ground wires that were connected using two wire nuts.  One of those black wires is also the antenna, it's the one that comes from a 2-pin white connector on the motherboard, the other pin will be unpopulated.  It was also connected with a wire nut to the handrail wires.  The wire from the trailing truck just connects the wheel wiper to the frame under a screw near the rear.

When in doubt, get your continuity meter and ID the antenna wire.  One probe on pin 23 (antenna) on the R2LC board and hit the suspect wire until you get continuity.  That antenna wire should go to your handrails and connect it to the handrail circuit with the original wire nut.

Next hit your rollers with your probe and find the (usually red) wire that shows you continuity.  Next hit your R2LC board pins 1 or 2 (hot) and identify that wire.  It should be coming from a white Leoco connector on the motherboard.   Lionel usually twisted those wires together and the others in that bundle should be to your smoke on/off switch and these are wire nutted.  On your driver board, identify the black ground wire.  Next to it will be the red (roller hot) wire.  These go into the black Molex plug.   If that goes to your motherboard, things will be connected to pins 1 and 2 on your R2LC.  On some that red wire if it is not hooked to anything will go to the red wire (hot) bundle.  The remaining wires will be ground.  Connect them together and check continuity with pins 3 and 4 on your R2LC board.  

Most of these connections are done with wire nuts.  Some and most today are just wrapped together with a chunk of shrink wrap on them.  Not a desired way to do it right.  Yours look like they were wire nutted.

The big thing you have to ID the antenna wire.  After that you will be fine.  

Last edited by Marty Fitzhenry

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