As mentioned, the coupler does not swing as smoothly. Also the tighter springs and looser specs cause the knuckle to be stiffer to open. Those are the two major characteristics which cause me to say they don't operate as smoothly.
However, they do work and stay coupled. I doubt they would hold for 1000 car train, but should hold just fine for anything under 100 cars - ie they should handle most any train on a home layout.
By the way, I about 95% successful in opening the Atlas boxes by using two small screwdriver blades to pull back the slotted tab at the rear of the box and release the bottom. Some come very easy others are harder, A very few break.