This is a non-modular RailSounds 2.5 item, so I'm not sure how you've re-seated the boards.
Try these:
Move the volume control pot up and down a few times to clear any oxidation.
If you have a PLCC chip extractor, pull the two chips from the RailSounds board, wipe with some contact cleaner, and reinstall.
Perform a restore/reset of the LCRU2 electronics in your unit using a "5."
TRW