Again, here is the problem. You have a TAS smoke unit and wiring setup for chuff input- I can see that in your wiring because I just worked on my own Weaver New Haven with the exact same smoke unit and wiring.
The point is, the 3 pin connector has smoke power (green), center pin (black) is frame ground, and the last wire is chuff (red wire and has a diode under heatshrink inline and visible in your photos.
However, when/if you plug in the Lionel standard 27 Ohm unit, the outer 2 pins are SHORTED together which potentially injects smoke AC power back down the chuff(red) wire)
Again this extremely bad and could damage the electronics of either the R4LC if they wired it for chuff input or the Railsounds board.
Either way, again, plugging that Lionel smoke unit in is bad in this wiring harness.
TAS turbosmoke instructions https://ogrforum.com/...20Turbosmoke%201.pdf
FYI, the reason I had mine open was my Weaver did not puff, smoke unit was constant on, but that was because the smoke unit jumper on the TAS unit was set to diesel mode. Changing that jumper again, was why I had mine open and how I have immediate understanding of your pictures and relevant wiring.
I cannot stress this enough, if/when you plug in that Lionel unit, it shorted AC hot smoke unit power into the chuff sensor/switch line- which is further connected to other electronics in the tender mostly 5V DC circuits.
This below is NOT compatible with this circuit as wired.