Here are my actual measurements on common Athearn-style HO wheels. I have bunches of each kind, just the wheels on axles without the truck assembly. These are spares acquired secondhand and I don't know what cars they were on originally. I've used some of these wheels to upgrade misc older HO cars.
smaller freight wheel tread dia 23/64" = 0.359" scales out to 31.3"
flange dia 55/128"= 0.430" scales out to 3.06" flange depth
larger freight wheel tread dia 3/8" = 0.375" scales out to 32.7"
flange dia 7/16" = 0.438" scales out to 2.72" flange depth
passenger wheel tread dia 51/128" = 0.398" scales out to 34.7"
I'm measuring with vernier calipers which are accurate to 1/128". The question is: why the slightly different sizes of freight wheels? On closer examination I discovered the flange depths and profiles are slightly different. I reckon Athearn modified the wheels to better conform to NMRA recommended standards at some point, maybe around 1970??
The difference in size between the smaller and larger freight wheels is so slight, you wouldn't notice it without measuring. The passenger wheels are noticeably larger, and also obviously different because they have one metal wheel and one plastic wheel on each axle.
I also have some of the larger freight wheels on brass axles, otherwise identical - who made them? Some folks wanted non-magnetic axles for easier operation with magnetic uncouplers.
MDC freight wheels appear identical to Athearn. My MDC cars from 1970's - 1980's have the slightly larger 0.375" wheels. I understand Cox was a rebranded Athearn product, maybe they had some variations on standard Athearn?
In the real world 33" wheel diameter is standard for most freight cars up to 70-ton capacity, 36" for 100-ton, 38" for 125-ton, 28" on some piggyback cars to lower the deck.
I've always liked the Athearn wheelsets because they have the reliable NMRA RP-25 wheel profile, they are uniformly round with smooth finish, the steel needlepoint axles run true and roll freely. Sometimes the wheel gauge needs slight adjustment to conform exactly to an NMRA wheel gauge.
I've encountered some of the supposedly better metal replacement wheels that had rough treads and flanges that were prone to derailment. And some metal wheels on plastic axles that weren't always true. I had bad results with Central Valley trucks [1970's] because the wheel treads had visible machining grooves which were derailment prone, even though the trucks were sprung and equalized. Athearn wheelsets were good performers and good value.
I run moderately long HO trains on a mountain layout with sharp curves and steep grades. Performance is reliable with good smooth wheels properly gauged. Vintage three-rail O-gauge is sometimes a challenge in this regard because of sloppy standards.
Anyone else out there who is interested in these details and has information to add?